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This is my first time encountering the strange marking under some vowels (i.e. e, i, a,). This symbol looks like a small comma underneath such vowels like in "*towi" under 102 and "nga(n)ɨɨ" under 105. See, they look like the underhook, but not exactly, so I'm not sure what symbol to transcribe that as. A note on the page indicates it as a "Intensifier".

The marks under the vowels are cedillas, and they mark nasalization of the vowel.